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COMPREHENSIVE INTERACTIVE REPORT · Debated grouping

Middle Pleistocene Homo

Heidelbergensis-grade populations773,000–300,000 years ago · Africa and Eurasia
Evidence-led reconstruction of Middle Pleistocene Homo
Scientific reconstruction · appearance remains interpretive · clothing: unknown

Current synthesis

A population-level framework replacing the assumption that every African and Eurasian fossil belongs to one uniform Homo heidelbergensis species.

01

Anatomy

Large brains and robust crania with major regional variation.

02

Ecology & food

Wide range of African and Eurasian habitats.

03

Technology

Acheulean and regional Middle Pleistocene technologies, wooden weapons and repeated-use sites.

04

Clothing evidence

Hide working occurs in parts of the Middle Pleistocene record, but species-level attribution and routine clothing remain uncertain. Reconstruction is shown without garments.

05

Behavior & social evidence

Cooperative large-game hunting and controlled fire are documented at some sites, not universally.

06

Relationships

Includes populations near the ancestry of sapiens, Neanderthals and Denisovans.

07

Controversies

Taxon boundaries, fossil allocation and the identity of the last common ancestral populations.

08

What reaches forward

The template report’s single-species model is preserved as a hypothesis and explicitly qualified.

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Legacy interactive report ↗Research update · 10.1038/s41586-025-09914-y ↗Research update · 10.1038/s41559-026-02983-w ↗Research update · 10.1016/j.jhevol.2026.103841 ↗
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